At some points it was mentioned that the regard of parents do play a role. The understanding problem I have there is, if the parental regard is to be seen like in steps so a nest1 cannot have something that is showing further back in history. Or is it only talking about "visible" results?

So when I have 2 meeroos and they make 5 nests, is each of the nests having both parental genes in it or is nest1 carrying less than nest5 because at the point nest1 was made the parents weren't able to reach back far enough to both pass down those genetics?